Chamorro, William, Diego Marín-Armijos, Valeria Granda, and Fernando Z. Vaz-de-Mello. “Checklist With a Key to Genera and Subgenera of Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) Present and Supposed for Ecuador”. Revista Colombiana de Entomología 44, no. 1 (August 5, 2018): 72–100. Accessed May 6, 2024. https://revistacolombianaentomologia.univalle.edu.co/index.php/SOCOLEN/article/view/6545.