CHAMORRO, William; MARÍN-ARMIJOS, Diego; GRANDA, Valeria; VAZ-DE-MELLO, Fernando Z. Checklist with a key to genera and subgenera of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) present and supposed for Ecuador. Revista Colombiana de Entomología, [S. l.], v. 44, n. 1, p. 72–100, 2018. DOI: 10.25100/socolen.v44i1.6545. Disponível em: https://revistacolombianaentomologia.univalle.edu.co/index.php/SOCOLEN/article/view/6545. Acesso em: 30 dec. 2025.